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Correspondence |

Evolution, Exaptation, and Stereopsis

Henrik Zetterberg, MD, PhD; Madeleine Zetterberg, MD, PhD
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Copyright 2005 American Medical Association. All Rights Reserved. Applicable FARS/DFARS Restrictions Apply to Government Use.

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Arch Ophthalmol. 2005;123(9):1281-1281. doi:10.1001/archopht.123.9.1281-a
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We read with great interest the article by Brodsky1 on the evolution of human stereopsis. Brodsky shows that the oblique muscles in humans seem to have abandoned their previous role in lower organisms in promoting panoramic vision and instead serve to stabilize the eyes to facilitate stereopsis. As Brodsky points out, this may be a very concrete example of evolutionary exaptation. This controversial term was originally coined by Gould and Vrba2 and means that a certain gene or trait that has evolved to fulfill a specific function takes on a new role (ex) and gets a new function (aptus) for which it was not originally designed.

We wish to report that accumulating data from functional analyses of homologous genes in an increasing number of different organisms support the view that exaptation indeed is an important mechanism during evolution. Again, an interesting example can be found in the evolutionary history of the eye. In the zebrafish embryo, the not really finished (nrf) gene is essential for the development of photoreceptors.3 However, its primary role in humans is to facilitate nuclear-mitochondrial interactions in a wide variety of cell types and in chickens to promote erythropoiesis.3 Similarly, the intraflagellar transport genes used by Chlamydomonas to make motile flagella are also used to assemble cilia in chemosensory neurons sensing salt concentration in Caenorhabditis elegans and in mechanosensitive kidney epithelial cells in vertebrates.4 These examples and the article by Brodsky show that the same genes (and muscles) can be used for very different tasks in different organisms and underscore the validity of the exaptation concept for the understanding of evolutionary processes.

AUTHOR INFORMATION

Correspondence: Dr H. Zetterberg, Center for Neurologic Diseases, Brigham and Women’s Hospital, Harvard Medical School, Harvard Institutes of Medicine, HIM-610, 77 Avenue Louis Pasteur, Boston, MA 02115 (hzetterberg@rics.bwh.harvard.edu).

REFERENCES

Brodsky  MC. Do you really need your oblique muscles? adaptations and exaptations. Arch Ophthalmol 2002;120820- 828
PubMed
Gould  SJ, Vrba  ES. Exaptation: a missing term in the science of form. Paleobiology 1982;84- 15
Becker  TS, Burgess  SM, Amsterdam  AH, Allende  ML, Hopkins  N. Not really finished is crucial for development of the zebrafish outer retina and encodes a transcription factor highly homologous to human Nuclear Respiratory Factor-1 and avian Initiation Binding Repressor. Development 1998;1254369- 4378
PubMed
Amsterdam  A, Nissen  RM, Sun  Z, Swindell  EC, Farrington  S, Hopkins  N. Identification of 315 genes essential for early zebrafish development. Proc Natl Acad Sci U S A 2004;10112792- 12797
PubMed

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Brodsky  MC. Do you really need your oblique muscles? adaptations and exaptations. Arch Ophthalmol 2002;120820- 828
PubMed
Gould  SJ, Vrba  ES. Exaptation: a missing term in the science of form. Paleobiology 1982;84- 15
Becker  TS, Burgess  SM, Amsterdam  AH, Allende  ML, Hopkins  N. Not really finished is crucial for development of the zebrafish outer retina and encodes a transcription factor highly homologous to human Nuclear Respiratory Factor-1 and avian Initiation Binding Repressor. Development 1998;1254369- 4378
PubMed
Amsterdam  A, Nissen  RM, Sun  Z, Swindell  EC, Farrington  S, Hopkins  N. Identification of 315 genes essential for early zebrafish development. Proc Natl Acad Sci U S A 2004;10112792- 12797
PubMed

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